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Through a series of misunderstandings, Alvin, Simon and Theodore come to believe that Dave is going to propose to his new girlfriend in Miami…and dump them. They have three days to get to him and stop the proposal, saving themselves not only from losing Dave but possibly from gaining a terrible stepbrother.
《鼠来宝4：萌在囧途》还是以三只可爱的花栗鼠为主角，讲述了花栗鼠艾文 （贾斯汀•朗 Justin Long 配音）、赛门（马修•格雷•古柏勒 Matthew Gray Gubler 配音）以及喜多（杰西•麦卡尼 Jesse McCartney 配音）由于一系列误会，以为它们的主人大卫（杰森•李 Jason Lee 饰）要在迈阿密向其新女朋友求婚，并有可能抛弃它们。它们得赶往迈阿密，找到大卫并阻止求婚。因为这样不仅不会失去大卫，还可能避免获得一位可怕的伙伴 ——大卫新女友的“小恶魔”儿子。一场跨越美国的音乐之旅即将爆笑启程。